4XXR series is different from the 5XX regular series in a few visible ways.. Well you can do this comparison with the 5XX and 5XXR
the "R" in the 4XX series stands for Reverse lip. This is a flat across dish which is polished and gives the rim a bigger appearance. It gives it a bigger appearance because the spoke can go all the way to the end of the rim
They are both 3 piece forged wheels as far as I know. 4 and 5 have different styles, and the 5XX series seem to "curve out" while the 4XX look more flat.
A standard 5XX has a bevelled "step" lip which doesn't go straight across. The spoke section won't be as big as a reverse lip wheel
I'm not 100% sure about the construction of the wheel. I learned this all before, but I forget details
There is a reason 5XX series is more expensive than 4XX series!
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